So, is this an optional, third-party theme? Or will this be how Firefox looks on GNOME by default?
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It's a third party theme. Firefox uses GTK as it's toolkit but almost all of the browser is styles through html/css, this theme replaces most of the browser with GTK styling instead.
Ah, now I gotta nitpick you on your Firefox facts.
Firefox mostly uses the XUL/XPCOM toolkit, which was written by Mozilla. It only references the GTK theme for how it should theme itself, and it makes use of the GTK file picker.
XUL code is rather similar to HTML, so one can also use CSS to style it, which is what this theme does to make it look more like a GNOME application.
Thanks for explaining, I wasn't sure how it worked inside but I thought I knew the basic idea.